Animal behaviour can be modified substantially through interaction with humans, whether non-lethal (e.g. coexistence, tourism viewing or feeding) or lethal (e.g. hunting. fishing, or collection). An extensive liter ature has documented these changes in many of the world's ecosystems (reviewed by Stankowich and Blumstein, 2005; Cooper and Blumstein 2015), and documents many similarities in altered behaviour of animals in terrestrial, freshwater, and marine environments. For example, the non-lethal presence of humans in terrestrial environments often results in increased flight distance for a variety of species, including birds and large-bodied ungulates (e.g. de Boer et al. 2004; Thiel et al., 2007) This trend of increased flight distance also occurs when humans hunt animals, often inducing greater changes in behaviour compared to non-lethal interactions with humans, regardless of the species or eco-system (Jayakody et al., 2008; Guidetti et al., 2008).