REPRODUCTIVE TIMING AND FREQUENCY
Four females were observed breeding in all three breeding
seasons, indicating that individual B. variegatus females
were capable of breeding annually (Fig. 2a). In addition,
B. variegatus females had reasonably distinct breeding seasons
where the median birth date was 24 February and
59% of births occurred in January, February and March
(n = 17; Fig. 3). Estimating the duration of juvenile dependence
was challenging, but one juvenile that was observed
on a nearly daily basis remained with its mother for
109 days. The proportion of adult females that produced
young from 2010 to 2012 was 0739 (34 of 46 breeding
attempts; SE = 0065) and varied little among years
(min = 0700, SE = 0145; max = 0778, SE = 0098).
Reproduction in C. hoffmanni was more asynchronous,
with young being born in all months except August
(Figs. 2b and 3). Moreover, C. hoffmanni young were
dependent on their mothers for longer than B. variegatus,
with the average duration of maternal dependence lasting
177 days (range = 139–208 days) among four young
tracked regularly from birth through independence. For
C. hoffmanni, the proportion of adult females that produced
young was 0868 (SE = 0055; 33 of 38) in 2010
and 0682 (SE = 0070; 30 of 44) in 2011. Overall, the
annual proportion of breeders was 0768 (SE = 0046; 63
of 82).