The chemical composition of SH and TSH was measured as follows. The lignin content was determined according to a standard method of Technical Association of Pulp and Paper Industry TAPPI T222 om-88. The holocellulose (α-cellulose + hemicellulose) content was estimated by the acid chlorite method (Browning, 1967). The α-cellulose content was determined treating the holocellulose with potassium hydroxide solutions (Browning, 1967). The hemicellulose content was found by subtracting the α-cellulose part from the holocellulose content. An average of three replicates was calculated for each sample.