The rotation of the earth causes sunlight levels to vary in predicable patterns of night and day with gradual increases and decreases in photosynthetically active radiation (PAR). Environmental variables like cloud cover, sunflecks, gradual vertical mixing in natural aquatic systems impose rapid changes in PAR that are superimposed on the aforementioned day/night cycles [1] and [2]. Turbulent mixing in turbid photobioreactors has similar effects. Understanding the physiological response to fluctuations in light fluxes is important for parameterizing models of photosynthetic productivity and for providing directions for improvements of overall biomass yields [3] and [4].