The nonparametric tests indicate that interaction between DA and sampling site on FA is not significant (Table 4).This suggests that variability among sampling sites in FA is mainly due to variability in DI rather than DA. On average, DA is marked bythe fact that columns IIa and IIIb are the shortest, with less primary tubercles across all sampling sites for P. lividus and higher number of pore pairs in A. lixula.