Acknowledgements The authors are indebted to S. Kwebiiha, E. Tukesiga, J. Mpagi and other staff members for their support during the field work and to P. Eggert and K. Pähle for their excellent technical assistance in the laboratory
AcknowledgementsThe authors are indebted to S. Kwebiiha,E. Tukesiga, J. Mpagi and other staff members fortheir support during the field work and to P. Eggertand K. Pähle for their excellent technical assistancein the laboratory