2.1. Yeast strains
Yeast strains collected and taxonomically identified by Takashima et al. were used as the main screening resource (Takashima et al., 2012). The yeast strains isolated from the campus of Kyoto University (Kyoto, Japan) were also assessed. As control strains, L. starkeyi NBRC 10381 and R. toruloides NBRC 0559 were obtained from the National Institute of Technology and Evaluation (NITE) Biological Resource Center.