Sampling was carried out at a jetty on the edge of the reef (depth: ca. ∼10 m) during four study periods; 22–24 August 2004, 1–3 October 2004, 25–27 February, and 2–4 June 2005. We therefore collected samples in multiple monsoonal periods from two SW monsoon periods (a late SW monsoon, August 2004 and anearly SW monsoon, June 2005), an inter-monsoon period (October 2004), and a NE monsoon period (February 2005).