f you leave or are kicked from the guild, your skills will be lost until you join another guild. Once the skills are restored, it will be same as your old guild's skill tower. So, if you have purchased level four in a guild skill from a previous guild, but your current guild has only purchased level two, your guild skills will be at lvl four.
VaultEdit
This is simply extra storage space.
Each upgrades increases the storage limit by 5 items.
When a player leaves a guild or is kicked, the items placed in the vault by that player will be sent to that player by the system in the mail.