This research was supported by Basic Science Research Program through the National Research Foundation of South Korea (NRF) funded by the Ministry of Education, Science and Technology (Grant number 2011-0001108), the Advanced Biomass R&D Center (ABC) of South Korea Grant funded by the Ministry of Education, Science and Technology (ABC-2011-0028387), Marine Biotechnol-ogy Program Funded by Ministry of Land, Transport and Maritime Affairs of South Korean Government, South Korea and the Man-power Development Program for Marine Energy funded by Minis-try of Land, Transportation and Maritime Affairs (MLTM) of South Korean government and by the World Class University (WCU) pro-gram through the National Research Foundation of South Korea funded by the Ministry of Education, Science and Technology