Similarly, when S. thermophilus LY03 and S. thermophilus Sfi20 are grown in a medium with both glucose (2.5% and 5.0%,
respectively) and lactose (5.0% and 2.5%, respectively), the HePS production is enhanced as compared to growth in media with glucose (7.5%) or lactose (7.5%) as the sole carbohydrate sources (Degeest & De Vuyst, 2000; Degeest et al., 2001b).