The original LAMTOR1 copy was expressed
in all bowhead whale adult tissues for which we have
data, with the duplicate having much lower (but detectable)
expression in heart and retina. Also of note, putative duplications
of 26S proteasome non-ATPase regulatory subunit 4 (PSMD4)
and ubiquitin carboxyl-terminal esterase L3 (UCHL3) were identified
with evidence of expression, which is intriguing considering
the known involvement of the proteasome-ubiquitin system in
aging (Lo´ pez-Otı´n et al., 2013)