farming on diversity or species richness. Our studies
contributed to the large heterogeneity among studies at
the landscape scale (Table 2). We conclude that positive
effects of organic farming on species richness and
diversity can be expected in intensively managed
agricultural landscapes, but not necessarily in smallscaled
mosaic landscapes with a mixture of agricultural
fields and non-cropped habitats.
Our meta-analysis shows that biodiversity can be
expected to benefit from organic agriculture. Additional
studies published after we completed our literature search
largely confirm this finding (Kremen, Williams & Thorp
2002; Aude, Tybirk & Bruus Pedersen 2003; Hutton &
Giller 2003; Hyvönen
et al
. 2003; Mulder et al. 2003),
although, as in our analysis, there are exceptions
farming on diversity or species richness. Our studiescontributed to the large heterogeneity among studies atthe landscape scale (Table 2). We conclude that positiveeffects of organic farming on species richness anddiversity can be expected in intensively managedagricultural landscapes, but not necessarily in smallscaledmosaic landscapes with a mixture of agriculturalfields and non-cropped habitats.Our meta-analysis shows that biodiversity can beexpected to benefit from organic agriculture. Additionalstudies published after we completed our literature searchlargely confirm this finding (Kremen, Williams & Thorp2002; Aude, Tybirk & Bruus Pedersen 2003; Hutton &Giller 2003; Hyvönenet al. 2003; Mulder et al. 2003),although, as in our analysis, there are exceptions
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