Guild structure of the dung beetle communities at Usina Serra Grande, north-eastern Brazil. Percentage of individuals (mean ± SE) (standardized to fragment size)
according to (A) diet and (B) resource-relocation behaviour; and percentage of body mass (mean ± SE) (standardized to fragment size) according to (C) diet and (D) resourcerelocation
behaviour between fragments, arranged in two size classes: small fragments (100 ha) (black bars).