In the conventional classification system, style length wasone of the major recognizable female traits used to identify thespecies (Bureau, 1873; Koidzume, 1917; Zhou and Gilbert, 2003;Chang, 2006b). This character serves to separate species depend-ing whether the style length exceeds 0.5 mm: the ‘Dolichostylae’(>0.5 mm style length) and ‘Macromorus’ (