Large-leafed species generally also possess large buds, flowers and fruit,
and stout twigs. Frequently, taxa differ in leaf and twig size, and occasionally
number of nerves, but share distinctive leaf and twig shape, indumentum, and
other characters in common; these are almost always allopatric and are
recognized as subspecies (10). Occasionally, as in Shorea macroptera (6)
subspecies may co-occur. In this example, no morphologically intermediate
individuals occur in the mixed stands, suggesting that even when distinctions
are reduced to this level of subtlety the taxa may function as biological
species.