However, the strictly aquatic water lily lineage (Nymphaeales) follows Amborella as the subsequent sister to all other flowering plants. As a result of the near-basal placement of Nymphaeaceae, the habit of the first flowering plants is reconstructed as equivocal on the basis of analyses of extant taxa (Soltis et al. 2005). Nonetheless, the early appearance of the water lily lineage, together with the discovery that some of the oldest angiosperm fossils are aquatic (e.g., Archaefructus), reinforced the hypothesis that the earliest angiosperms may have been aquatic—that is, “wet and wild” (D. Dilcher, pers. comm.; see Coiffard et al.2007).