At saltmarsh, temperature did not change significantly during the incubation (mean: 18.6 °C; SD: 0.9 °C). In this zone, bioturbation enhanced the CO2 flux (ANOVA: F2,15 = 80.48, P< 0.05; Fig. 5e), whereas there was no bioturbation effect on benthic fluxes of both carbohydrates and proteins (Fig. 5f and g). DOC flux was toward the sediment at non-bioturbation treatment, but it was toward the water column at both bioturbation treatments (i.e., B and B + C; Fig. 5h).