Fragment surveys included trees with a DBH > 10 cm (diameter at breast height), which were classified with regard to regeneration strategy (pioneer, and shade-tolerant
trees) and forest vertical stratification (understory, canopy,and emergent trees).
In this study, tree density (per plot), tree richness (per plot) and percentage of shade-tolerant species (i.e., those capable of regenerating in the shaded understorey of old growth
forests) and emergent species (i.e., species capable to occupy the highest stratum of the forest) were used as tree community attributes