Pelophylax perezi
(Lo´pez Seoane, 1865), Pelobates cultripes (Cuvier,
1829), and Pleurodeles waltl Michahelles, 1830 were
the amphibian species with the highest co-occurrence
with P. clarkii. During the pre-drought period, crayfish
co-occurred with most amphibian species in different
areas of the park, except in the southern ponds.