whereas no such change was
observed under the dark in in vitro fruits or LL in in vivo fruits. Only in in vitro fruits under HL, activities of
the antioxidant enzymes such as ascorbate peroxidase, dehydroascorbate reductase, and catalase were
higher in salt-stressed fruits than in controls. In contrast, oxidative parameters on a DWbasis, as indicated
by malondialdehyde, remained unchanged in both in vitro and in vivo fruits. In a principal component
analysis, salt-induced changes in antioxidant systems showed clear differences between in vitro and
in vivo fruits and between light conditions. Thus, we conclude that salt-induced changes in the ASA
content and activities of antioxidant enzymes cannot be accounted for by salt stress alone and that high
light intensity is also an important factor inducing the upregulation of antioxidant enzymes that reduce
salt-induced oxidative stress.