Macrolecithal[edit]
A baby tortoise begins to emerge "fully developed" from its macrolecithal egg.
Eggs with a large yolk are called macrolecithal. The eggs are usually few in number, and the embryos have enough food to go through full fetal development in most groups.[7] Macrolecithal eggs are only found in selected representatives of two groups: Cephalopods and vertebrates.[7][16]
Macrolecithal eggs go through a different type of development than other eggs. Due to the large size of the yolk, the cell division can not split up the yolk mass. The fetus instead develops as a plate-like structure on top of the yolk mass, and only envelopes it at a later stage.[7] A portion of the yolk mass is still present as an external or semi-external yolk sac at hatching in many groups. This form of fetal development is common in bony fish, even though their eggs can be quite small. Despite their macrolecithal structure, the small size of the eggs does not allow for direct development, and the eggs hatch to a larval stage ("fry"). In terrestrial animals with macrolecithal eggs, the large volume to surface ratio necessitates structures to aid in transport of oxygen and carbon dioxide, and for storage of waste products so that the embryo does not suffocate or get poisoned from its own waste while inside the egg, see amniote.[9]
In addition to bony fish and cephalopods, macrolecithal eggs are found in cartilaginous fish, reptiles, birds and monotreme mammals.[3] The eggs of the coelacanths can reach a size of 9 cm in diameter, and the young go through full development while in the uterus, living on the copious yolk.[17]