Augustine grass at the Crago Sod Farm (South Bay, Florida) as being closest to H. wessoni. Ye & Robbins (2000) examined several populations of H. wessoni from different geographical origins, compared them with the type specimens of H. annulatus, and found that the small differences observed in H. annulatus were within the intraspecific variation of H. wessoni, thus supporting the synonymy of these two species.