Brümmer et al. (2012) reported that althoughclimate conditions, LAIs and ecosystem biomasses of grassland,peatland and forest differ, their ecosystem-scale WUE values arewithin a relatively stable constant range (2.6–3.6 g CO2kg−1H2O).Orchard-scale WUE is also influenced by soil moisture content;for example, De Azevedo et al. (2006) found that WUE decreasedas water irrigation increased in a 6-year-old coconut tree plan-tation.