However, the use of only mtDNA as a genetic marker in molecular ecology and phylogeographic studies possibly leads to misinterpretation and may yield an incomplete evolutionary history of the target species [28,29]. This may be due to mitochondrial recombination in population levels [30], mitochondrial introgression [31-33], and the indirect selection caused by symbionts [34]. Thus, the combined used of mitochondrial and nuclear DNA, as opposed to the sole usage of mtDNA markers, could allow researchers to answer more complicated evolutionary questions about target organisms