Anyhow, the last three or six segments of the antennal club of some are passalids are brought closely together by partial rolling up of anntenae.
Represented by seven subfamilies, totally 140 species are illustrated in this book. They are , Dynastinae 17, Euchirinae 2, Copriinae 20, Geotrupinae 1 , Melolonthinae 2 ,Rutellinae 16 and Cetoninae 82 species