In addition to being effected by many environmental factors, the expression levels of antho-cyanins biosynthetic genes are also modulated by transcriptionfactors (TFs), such as ELONGATED HYPOCOTYL5 (HY5), basic helix-loop-helix (bhlh) and v-myb avian myeloblastosis viral oncogenehomologue (MYB) , which bind tostructural gene promoters to modulate gene expression