Dragonflies are accomplished aerial pursuit hunters: they can hover, accelerate in almost any direction and manoeuvre precisely at high to intercept other insects, with measured success rates as high as 97%. The direct flight musculature of dragonflies means that stroke frequency, amplitude, phase and angle of attack can be varied independently on each of the four wings. Dragonflies put these abilities to excellent effect, using them to enlist a variety of wing . The wings counterstroke when cruising, but stroke in-phase during manoeuvres and when higher accelerations are required.