Egg shell strength was evaluated using a press meter (FHK, Japan) to gauge the force producing cracking under longitudinal compression, and the shell maximum deformation was recorded (g/cm2). The average shell thickness was measured on the large end, equatorial region and small end using a dial pipe gauge (Ozaki MFG. Co., Ltd., Japan). Finally, Egg weight, egg-yolk height,egg-yolk color, and Haugh units were evaluated using an egg multi-tester (Touhoku Rhythm Co., Ltd., Tokyo, Japan).