DNA–DNA hybridization was carried out between strain CNU040T and A. terrigena KCTC 12545T using the membrane filter hybridization method described by Brown (2005). Probe labelling and hybridization were conducted using the non-radioactive DIG-High Prime system and the hybridized DNA was visualized using the DIG luminescent detection kit (Roche), according to the manufacturer’s instructions. DNA–DNA relatedness was quantified using a densitometer (Bio-Rad).