Average discharge values (average ± SE) in the upstream section were 69.4 ± 1.9 m3s−1 during dry periods, and 275.6 ± 8.7 m3s−1 during wet periods. Average discharge values in the downstream section were173.3±2.9m3s−1 during dry periods, and 351.4 ± 8.1 m3s−1 during wet periods. Therefore, discharge was 1.5 times greater downstream than upstream of the reservoirs, but the generalflow patterns were similar in the two sections (Fig. 1). Dry periods had higher water temperature and conductivity than wet periods. Upstream sites had higher suspended solids and nitrate and ammonium concentration than downstream sites (Table 1)