All
isolates produce strong positive reactions for alkaline phosphatase,
leucine arylamidase, acid phosphatase, naphthol-
AS-BI-phosphohydrolase and b-galactosidase. Strong or
weak positive reactions are produced for esterase, esterase
lipase and valine arylamidase. The main quinones are
ubiquinone-8, menaquinone-8 and demethylmenaquinone-
8; ubiquinone-7 and -9 are also present.