Dispersion values in observed and null distribution were used with standardize effect size (SES) of the trait dispersion metrics relative to the null distribution communities. These value provided signature of the trait clustering or evenness. Positives value for SES/MPD and SES/MNTD indicate trait clustering witch suggest that habitat filtering was a significant factor governing community structure. Negative value of the SES/MPD and SES/MNTD indicate evenness, which suggest that competition was a significant factor.