Skeleton feeding of a leaf is defined as scraping out the leaf tissue between the veins by insects (McDonald et al., 2007). It is difficult to separate skeleton feeding and natural decomposition especially in fossil leaves. If the remains of the fossil leaf are still intact and the tissue is missing in only in few areas, then it is a skeleton trace fossil (Scott, 1992). The skeletonized areas are elongated, rectangular, or irregular in shape (Fig. 7A, B, D). Irregular skeleton feeding are observed on the extant leaves of Actinodaphne (Fig. 8B). Removal of inter-venal tissue on the surface of the leaf is also observed in some specimens.