on macrophytes and algae, plankton-feeder and
scavenger like tilapia. Tilapia can also partly rely
on periphyton grown on rice plants as well as
known to partly feed on macrophytes (Rothuis
et al. 1998). Furthermore, results from this study
strongly suggest that the benefit from integrated
rice–fish culture crucially depends on the fish harvesting
weight and rice production as well.