Ground beetle data from two Scottish ECN sites, Glensaugh and
Sourhope, were used for the analysis. Glensaugh Research Station
is located in North-East Scotland, 56 km south-west of Aberdeen
(latitude/longitude: 56.895363, −2.541496, altitude: 165 m, mean
annual temperature: 7.8 ◦C, mean annual rainfall: 1048 mm).
Sourhope lies 24 km south-east of Kelso (latitude/longitude:
55.481188, −2.245039, altitude: 231 m, mean annual temperature:
7.5 ◦C, mean annual rainfall: 893 mm)