Water stress is one of the most important environmental factors inhibiting photosynthesis [32]. Photosynthetically derived ATP and NADPH are transferred to a large number of biosynthetic reactions, including the reactions of the carbon fixation pathway. Carbon from CO2 enters the pathway and combines with ribulose 1, 5 biphosphate under the catalytic action of ribulose biphosphate carboxylase (rubisco). Subsequently, ribulose 1, 5 biphosphate is converted to triose phosphates, which are exported to the cell cytosol to provide an organic carbon source for multiple cellular reactions [4]. Many studies have shown that decreases in photosynthesis resulting from water stress can be associated with the perturbation of the biochemical processes of that pathway. Several in vivo studies demonstrated that water stress resulted in damage to the oxygen evolving