Thus, the correlation between microbial biomass and macroaggregate content could be due to the fungal biomass which may not have been directly related to the fungal hyphae length. No conclusion could be drawn about the higher fungal hyphae length in the control soil. However, it is possible that the endogenous recalcitrant substrates in the soil could increase the fungal biomass. Consequently, the presence of fungal hyphae could provide physical binding for particles as previously demonstrated but it could not wholly explain the formation and stabilization of aggregates [31]. In addition, it is known that roots and hyphae act together as binding agents between particles [32]. In this study, root biomass was correlated with the macroaggregate content and stability (MWD) whereas the fungal hyphae length was not.