Gentiana triflora and G. scabra are protandrous. Their
pistils are immature (top of stigmas are closed) when their
anthers are dehisced, and then the pistils gain the ability to
accept pollens when their stigmas are opened. The flower
bud stages prior to opening of the top of the stigma were
classified into six stages by three traits, length of flower
bud, anther dehiscence and the position between stigma
and anthers (Fig. 1)