AcknowledgementsThis work was supported by the National Natural Science Foun-dation of China (No. 31101364), the Ministry of Agriculture ofChina for Financial Support (No. CARS-25), the Special Fund forAgro-scientific Research in the Public Interest (No. 201203095),the Young Investigator Fund of Beijing Academy of Agriculturaland Forestry Sciences (No. 201404), the National Natural ScienceFoundation of Beijing (No. 6144020), Innovation Fund of BeijingAcademy of Agricultural and Forestry Sciences