Such studies are also rare on other continents where
these species occur. The literature shows only general
characterization, as the brief anatomical description
of the leaves and stems of Scutellaria pinnatifida A.
Hamit subsp. pichleri occurring in Iran (Hatamneia
et al., 2008), and a morpho-anatomical study of the
vegetative and reproductive organs of Scutellaria
orientalis L. subsp. bicolor (Hochst.) Edmund, subsp
santolinoides (Hausskn ex Bornm) (Ozdemir & Altan,
2005) and subsp. pinnatifida Edmondtson (Candan &
Çali, 2012), occurring in Turkey. The mai characteristics
described for Scutellaria are the presence of specialized
cells that have “scutellarin” on the abaxial epidermis;
diferents types of glandular trichomes which can be
a large single-cell, two-cells or more than sixteen
cells on the head and they can be penduculate or not.
These characteristics are emphatazing as important
to taxonomy and were used in identification keys to
distinguish this genus among the Lamiaceae (Metcalfe