However they were separated based on subtle morphological characters: B. papayae was distinguishable from B. dorsalis s.s. in having a longer aculeus and was deemed separate from B. philippinensis based on morphological variation in the scales on the distal end of the middle segment of the aculeus; and while B. philippinensis was recognised as ‘difficult to separate’ from B. dorsalis s.s. it was considered a new species based on differences in the mean ratio of wing vein lengths (CuA1 along dm cell) to the length of the aculeus (1.19 in B. philippinensis and 1.47 in B. dorsalis s.s.) [4]