Robbrecht & Puff (1986) published a comprehensive morphological study of Gardenieae based on several Rubiaceae systematic treatments by Schumann (1891), Verdcourt (1958), Bremekamp (1966), and their own observations and proposed a new classification (Table 1). Tribe Gardenieae was subdivided into two groups, subtribe Diplosporinae with ca. 20 gen- era and subtribe Gardeniinae including ca. 60 genera. Tribe Aulacocalyceae was described by Robbrecht & Puff (1986) to accommodate several genera previously placed in Gardenieae