The biogenic amines production was studied in Lactobacillus strains from fruits [49]. Lactobacillus
strains are considered to be generally safe. However the authors demonstrate that a strain of L. hilgardii
isolated from wine was able to produce agmatine and putrescine from one of the major amino acids
found in fruit juices and wine, arginine and its amino acid–derived ornithine.
Biogenic amines like tyramine as well as diamines such as putrescine and cadaverine have been
described as precursors of carcinogenic nitrosamines [50]. Alcohol may enhance the effect of amines