Tomato (Lycopersicon esculentum) is one of the world’s major vegetables with a worldwide production of 126 million tons in 2005 (FAOSTAT, 2007). It is an excellent source of many nutrients and secondary metabolites that are important for human health: mineral matter, vitamins C and E, b-carotene, lycopene, flavonoids, organic acids, phenolics and chlorophyll (Giovanelli & Paradiso, 2002).