This strain was identified by its morphological, physiological, and chemotaxonomic features and confirmed using a 16S rRNA-based phylogenetic analysis (Zheng et al., 2006). Bacteria cells were prepared by growing cultures for 48 h in a liquid medium that contained 10 g/L soluble starch, 5 g/L beef extract, 3 g/L peptone, 10 g/L sucrose, 5 g/L yeast extract and 5 g/L CaCl2 and had a pH between 7.2 and 7.4.