The abdomen of the female became swollen prior to spawning.
Spawning took place mostly during the morning; between
09:00 and 13:00 h. Females laid capsule-shaped eggs on the
cleaned substratum in almost circular or oval patches and subsequently
the male fertilized the eggs. The spawning activity
lasted from an hour to an hour and forty five minutes. The eggs
are adhesive and are covered with a transparent chorion and a
narrow perivitalline space. The eggs measured 2.0–2.1 mm in
length and 0.9–1.0 mm in width. Approximately 300–400 eggs
were spawned at a time. The number of eggs gradually increased
in subsequent spawning. One end of the egg capsule identified as
the animal pole contained some gelatinous substance to adhere
itself to the substratum.