Acknowledgements
We would like to thank the landholders and program adminis-
trators involved in this research and field assistants N. Browne and
C. Langbein. Thanks also to thosewho provided valuable comments
on drafts of this manuscript, in particular D. Marsh, T. Brewer,
N.Marshall, and P. Valentine, and to E.J.I. Lédéewho prepared Fig. 1.
Thank you also to two anonymous reviewers who provided
very valuable comments on an earlier draft of the manuscript.
This research was made possible with funding from James Cook
University