They show a considerable degree of diversification within Agaricomycotina. In our phylogenetic analysis, four clusters with high bootstrap support could be recognised within the group. The highest diversity of its members is observed in the crown groups of Agaricomycotina, i.e. orders of Agaricales and Boletales, whereas the diversity is lower in basal lineages (like Sebacinales and Cantharellales).