The physiological state of the bryophyte, and hence
desiccation tolerance, varies with the season. Most
mosses seem to be most sensitive during autumn and early
winter, the season when most bryophytes resume growth
(Dilks & Proctor 1976a). Tolerance increases from spring
to a maximum in early summer, the season when many
species become dormant. Some degree of acclimation may
be occurring, resulting in increased tolerance as summer
approaches (Richardson 1981).